To clairify further:
Both the intake and exhaust "air" flow is not "smooth". If you think about how the flow is produced you'll understand.
Air in drawn in only on each intake stroke of a piston. Therefore there is a regular pulse frequency associated with the intake air at any given engine RPM. There is also some relatively small power used in drawing in the intake charge.
Now here's how a tuned intake (and exhaust for that matter) works. If you adjust the intake chambers such that it is a resonant frequency of the intake flow at the desired engine RPM then you will see an increse in power / torque & MPG (as a consequence) since then pulses are now working together to improve overall flow. Again at the _designed_ RPM.
Honda has been doing this now for over 10 years with many of their models. The overall percentage gains are small, low single digits, and with MPG probably not discernable except under labratory conditions. But its essentially "free" for the taking. The've even done a duel stage tuned system whereby a secondary chamber was switched open & closed based on engine RPM to further improve the effect.
And just like as with the lean burn window if you drive outside the "box" you won't "see" it. An aftermarket high flow filter _will_ improve peak HP, at the high end of the tach. The "gain" will also be too small to "feel".
HTH!